PhD Biology, University of Washington, 2016

BSc Marine Biology, University of California Santa Cruz, 2001

AA Spanish, Cabrillo College 1998



H.A. Hayford, M. J. O’Donnell, E. Carrington (In review) Radio tracking detects thermoregulation at a snail’s pace.

W.B. Stickle, E. Carrington, H. Hayford (2017) Seasonal changes in the thermal regime and gastropod tolerance to temperature and desiccation stress in the rocky intertidal zone at the southern end of the inside passage. Journal of Experimental Marine Biology and Ecology, 488:83-91 Abstract

S.E. Gilman, H. Hayford, C. Craig, E. Carrington (2015) Body temperatures of an intertidal barnacle and two whelk predators in relation to shore height, solar aspect, and microhabitat. Marine Ecology Progress Series, 536:77-88 Abstract

H.A. Hayford, S.E. Gilman, E. Carrington (2015) Foraging behavior minimizes heat exposure in a complex thermal landscape. Marine Ecology Progress Series, 518:165-175 Abstract



experimental ecology – natural history – coastal communities

ecophysiology – thermal biology – behavior

climate change – environmental monitoring – ocean acidification

In my doctoral research, I found that the intertidal whelk snail, Nucella ostrina, moderates its exposure to high temperatures (thermoregulates) by feeding on reliably cool days of the two-week tidal cycle. I tested the effect of temperature on feeding preference behavior in experimentally created habitats in the field, used radio frequency identification (RFID) tags to track snail movements in their natural habitat, estimated snail body temperatures in microhabitats using physical thermal models, and assessed thermal performance as recovery time after stress.

>100 marine field sites in 3 countries (Canada, Chile, USA)

1000s of field days include rocky shores, sandy shores, estuaries, sloughs, docks, jetties, boats, lakes, streams, forests

Research Assistant, Partnership for Interdisciplinary Studies of Coastal Oceans (PISCO), University of California, Santa Cruz, 1999-2009



I have been fortunate to teach students from 2 years to over 80 in an equally wide variety of topics and settings: biology, marine science, journalism, roller skating, classrooms, labs, aquariums, and sports facilities. My teaching methods have developed from the similarities across these experiences. I prioritize establishing an environment of mutual respect and collaboration. I aim for students to describe patterns both verbally and mathematically, apply logic to propose mechanisms, and draw on knowledge from multiple disciplines to suggest solutions to problems. I try especially to foster appreciation for: natural history, the inevitable failures that come with the scientific process, and the importance of a diversity of participants.

Instructor of record for Marine Ecological Processes (University of Washington, Seattle)

Teaching assistant for 22 classes, including Marine Ecology, Ecology, Limnology, Invertebrate Zoology, Marine Algae, Physiology, Marine Research Apprenticeships, Cell & Molecular Lab Techniques, Marine Biotechnology, Introductory Ecology & Evolution, Introductory Cell & Molecular Biology (University of Washington, Seattle & San Jose State)

Ingrith Deyrup Distinguished Teaching Award (University of Washington, Seattle)






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